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Most infamously, Angron ordered that the World Eaters conquer their targeted worlds within 31 hours - a single Nucerian day and the time he had scored his greatest victory upon the planet. The World Eaters consistently failed to capture worlds within this time limit, and each time Angron ordered decimation as punishment, forcing 1 of every 10 World Eaters to be killed by his other 9 brothers. After several of these episodes, Centurion [[Mago]] finally refused to again enact decimation after the [[Ghenna Massacre|Ghenna Campaign]]. This caused Angron to be thrown into a raged frenzy, during which he murdered several nearby World Eaters until being subdued by his [[Librarian]]s.{{Fn|25b}} While Angron lay comatose and had his memories of Nuceria absorbed by the young [[Lexicanium]] [[Tethys]], a crisis erupted across the Legion as [[Mago]] attempted to stop [[Gahlan Surlak]] from installing the perfected Butcher's Nails in all of the World Eaters. Mago's revolt came to its climax on [[Ghenna]], where Kharn slew the rebel [[Centurion (Rank)|Centurion]] while Angron slew Tethys.{{Fn|25f}} Now at the head of a Legion wielding the Nails, Angron reaped many victories during the [[Great Crusade]], although some criticized the extreme and bloodthirsty tactics he used to ensure the destruction of his opponents.{{Cite This}}

At some point following the rediscovery of Angron, [[Arkhan Land]] was brought to a secret laboratory hidden in a dormant volcano in tundra location on Terra to aid in the Emperor's investigation of Angron's crude augmetics as the primarch lay unconscious. Land had previous experience with the device taking over Angron's body, having see it when investigating the Hexarchion Vaults before he ordered them sealed do to the threat the items within presented to the Machine Cult. He told the Emperor that it was called a ''cruciamen'', but the versions he saw were more crude. The Emperor explained that the implants were not his doing and that he believed they led to the Angron's instability. He also explained that they were recreated to remove the ability to enjoy anything but anger. When asked if the Emperor could remove it, he admitted he could, but the tendrils were deep in Angron's head and spine and he was missing his limbic lobe and insular cortex. The Butcher's Nails therefore replaced part of Angron's brain, allowing him to live.{{fn|19}}

*{{Endn|28}}: [[Saturnine (Novel)]] Part 2, Chapter 6

**{{Endn|28a}}: Part 3, Chapter 5

{{Primarchs}}